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There is currently no route to the mailbox database after DAG server failure

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This is an environment with Exchange 2007 and Exchange 2013.  Email is currently flowing through 2007 but all mailboxes reside in 2013.  There is a DAG setup in 2013.  We were performing HA testing and noticed when we brought down the Exchange 2013 DAG member holding the primary active manager mail flow stopped between 2007 and 2013.  Email just stayed in the queue and finally went to a queue with the error in the title.  To resolve we had to bring the server back online and restart the hub transport service on 2007 box.  Any ideas why this is happening?  Thanks.

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