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Email flow problem between two site

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We've configure 2 different AD sites and both the site are configured with Ex2010 SP1 server with different MailboxDB. We are facing problem while trying to send/receive internal email between two sites also cant receive email from the external.

Site-1: CAS + HT + MB (Mailboxdb1)

Site-2: CAS + HT + MB (Mailboxdb2)

Site-1 : Can send and receive mail to the external as well as the Mailboxdb1 but cant send to the Mailboxdb2

Site-2 : Mailboxdb2 users cant send/receive mail to Mailboxdb1 users but can send mail to the external only.

ERROR: There is currently no route to the mailbox database.

Please advise


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