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Hybrid Mail Flow

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There is a hybrid environment with exchange 2010.

Some users mailboxes on exchnage online, some others in on-prem exchange.

My question is;

when a user from external domain send mail to my  domain where mailbox is on exchange online not in on-prem.

--mail comes my firewall to check this user on my on-prem or not ...and then ? if not ? how this flow works ?


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