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External users unable to email distribution lists with error 550 5.1.1 User unknown

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We've recently run in to a problem with some external users trying to email some of our distribution lists. The following bounce message is sent to them:

This message was created automatically by mail delivery
software.

A message that you sent could not be delivered to one or
more of its recipients. This is a permanent error. The following address(es)
failed:

  distributionlist1@domain.com    host smtpserver.domain.com [xxx.xxx.xxx.xxx]    SMTP error from remote mail server after pipelined sending data block:    550 5.1.1 User unknown  distributionlist2@domain.com    host smtpserver.domain.com [xxx.xxx.xxx.xxx]    SMTP error from remote mail server after pipelined sending data block:    550 5.1.1 User unknown  distributionlist3@domain.com    host smtpserver.domain.com [xxx.xxx.xxx.xxx]    SMTP error from remote mail server after pipelined sending data block:    550 5.1.1 User unknown

The Default Frontend Transport Service logs show the following:

rcpt to:<distributionlist3@domain.com>
250 2.1.5 Recipient OK
DATA
354 Start mail input; end with <CRLF>.<CRLF>

Tarpit for '0.00:00:05' due to '550 5.1.1 User unknown'
550 5.1.1 User unknown

Any ideas?


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