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Unable to send mail from webmail to exchange server 2010

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Am deploying  Ms Exchange Server 2010, but I need you to assist me  to solve a problem.
This is the scenario below:
My client is made up three subsidiaries using same web mail service hosted by a provider.
One of the subsidiaries has decided to deploy their email in-house using Exchange Server 2010. So, while we are expected to migrate these  users to the Exchange server, remaining users from the two subsidiary will still be retained on  hosted web mail.
The users are to retain their email addresses in the new environment.
So far,
I have deployed Exchange Server, exchanging mail within the local network and outside domain like  yahoo.com, gmail.com etc.  This is working fine... OWA is also working fine.
Technical details:
Port 25 enabled on the firewall, 
External registered domain: tsllimited.com
Host name : mail.tsllimited.com
local domain:tsl.local
ISP has created MX record to tsllimited.
Since users are expected to retain their email addresses we configured  REPLY AS  to use   tsllimited.com (eg user1@tsllimited.com) so users are currently using same address format  as they have on the web mail.
But there is a problem, that is :
We can send mail to the hosted web mail  from the exchange server , but receivers can not deliver mail to the Exchange server from the web mail.
What do we need to solve this problem. PLS  HELP!!!!

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